Why a Floating Leaf Puts Its Stomata Upward

A water lily’s leaf has to exchange gases with the air while resting on the water. Its stomata—tiny adjustable openings in the leaf’s surface—are concentrated on the upper side, exposed to the atmosphere. Carbon dioxide enters through them for photosynthesis, while oxygen produced during that process can diffuse back out. Water vapor also escapes, a process called transpiration.

On a leaf held above land, stomata commonly occur on the lower surface, where they are less exposed to direct sun and drying air. A floating leaf faces the opposite problem: its lower side is pressed against water, so openings there would not have ready access to the air. Placing them on top makes gas exchange possible without lifting the leaf off the surface.

The Leaf’s Position Shapes Its Design

This arrangement works because a lily pad spreads across the water like a platform. Its broad blade reaches sunlight, while the stalk connects it to the plant below. The leaf’s upper surface is therefore both its main route for exchanging gases with the atmosphere and the side exposed to rain and splashes.

The adaptation is a useful reminder that leaves are shaped by where they sit, not just by what they do. A floating leaf and a land leaf can perform the same basic tasks—photosynthesis and gas exchange—but place their tiny openings on different sides to meet very different surroundings.